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A307941 Number of evolutionary duplication-loss-histories of the complete binary species tree with 4 leaves. 2
4, 34, 368, 4685, 66416, 1013268, 16279788, 271594611, 4660794200, 81747301898, 1458812278424, 26400987754054, 483374731032868, 8936983620559660, 166617056922535080, 3128790129161470470, 59124052722375912960, 1123458655726125274620, 21452847767668402271220 (list; graph; refs; listen; history; text; internal format)



An evolutionary history of size n is an ordered rooted (incomplete) binary tree with n leaves describing the evolution of a gene family of a species in phylogenomics. The complete binary species tree S of size k is a complete binary tree with k leaves. Any node of the history is associated to a unique node of S, where specifically every leaf is associated to a leaf of S. A history is created by the following process (note that intermediate trees in this process may not be valid histories): Start with a root node associated to the root of S. For a given tree in the growth process, choose a leaf and perform a duplication, speciation, or (speciation-)loss event. A duplication event creates two children both associated to the same node as its parent. A speciation or (speciation-)loss event can only occur if the node is associated to an internal node in S. In that case, a speciation event creates two children associated to the children of the node in S. A (speciation-)loss event creates only a left or right child, associated to the left or right child in S, respectively.


Table of n, a(n) for n=1..19.


G.f.: 1/2-(1/2)*sqrt(1+6*sqrt(-5+6*sqrt(1-4*z)+4*z)-6*sqrt(1-4*z)-4*z).


The complete binary species tree with 4 leaves is equal to


/ \

b c

/ \ / \

1 2 3 4

For convenience the internal nodes are labeled by a,b,c and the leaves by 1,2,3,4. The associated nodes in the histories will be denoted by the same labels.

The a(1)=4 histories with n=1 leaf are created by the following growth process:

a a a a

/ / \ \

b b c c

/ \ / \

1 2 3 4

after two loss events each.


(PARI) z='z+O('z^20); Vec(1/2-(1/2)*sqrt(1+6*sqrt(-5+6*sqrt(1-4*z)+4*z)-6*sqrt(1-4*z)-4*z)) \\ Jianing Song, Jul 29 2019


Cf. A000108 (caterpillar/complete binary species tree with 1 leaf, ordinary binary trees).

Cf. A307696, A307697, A307698, A307700 (caterpillar species tree with 2, 3, 4, 5 leaves).

Sequence in context: A274344 A199752 A264607 * A084973 A234313 A197921

Adjacent sequences: A307938 A307939 A307940 * A307942 A307943 A307944




Cedric Chauve, May 07 2019



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Last modified March 29 12:12 EDT 2023. Contains 361599 sequences. (Running on oeis4.)